Handbook of the Mammals of the World – Volume 9

Chief Editors:

Dr D. E. Wilson: Curator Emeritus, National Museum of Natural History, Smithsonian Institution, Washington, DC, USA.

Dr R. A. Mittermeier: Chief Conservation Officer, Global Wildlife Conservation; Honorary Member, International Union for Conservation of Nature (IUCN); and Chair, IUCN/SSC Primate Specialist Group, USA.

Authors:

Dr L. F. Aguirre: Researcher, Centro de Biodiversidad y Genética, Facultad de Ciencias y Tecnología, Universidad Mayor de San Simón, Cochabamba, Bolivia.

Dr F. C. Almeida: National Researcher, CONICET, Instituto de Ecología, Genética y Evolución de Buenos Aires, and Universidad de Buenos Aires, Buenos Aires, Argentina.

Dr J. Arroyo-Cabrales: Senior Scientist, Laboratorio de Arqueozoología “M. en C. Ticul Álvarez Solórzano”, Subdirección de Laboratorios y Apoyo Académico, Instituto Nacional de Antropología e Historia, Ciudad de México, Mexico.

Dr P. Benda: Research Curator, Department of Zoology, National Museum of Natural History, Prague; and Associate Professor, Department of Zoology, Charles University, Prague, Czech Republic.

Dr F. J. Bonaccorso: Emeritus Wildlife Biologist, U. S. Geological Survey, Hawaii Volcanoes National Park, Hawaii, USA.

C. J. Burgin: Researcher, Department of Biological Sciences, Boise State University, Boise, Idaho, USA.

S. F. Burneo: Mammalogy Curator, Museo de Zoología, Pontificia Universidad Católica del Ecuador, Quito, Ecuador.

M. A. Camacho: Associate Professor and Researcher, Sección de Mastozoología, Museo de Zoología, Pontificia Universidad Católica del Ecuador, Quito, Ecuador.

V. C. Cláudio: PhD Candidate, Instituto de Biologia, Universidade Federal do Rio de Janeiro, Rio de Janeiro, Brazil.

Dr G. Csorba: Deputy General Director, Hungarian Natural History Museum, Budapest, Hungary.

Dr E. R. Dumont: Dean, School of Natural Sciences, University of California, Merced, California, USA.

D. V. Espinosa-Martínez: Research Assistant, Department of Biology, Instituto Nacional de Antropología e Historia, Ciudad de México, Mexico.

Dr C. M. Francis: Manager, Wildlife Monitoring and Assessment, Canadian Wildlife Service, Environment and Climate Change Canada, Ottawa, Ontario, Canada.

G. S. T. Garbino: PhD Candidate, Departamento de Zoologia, Universidade Federal de Minas Gerais, Belo Horizonte, Brazil.

Dr N. P. Giannini: Principal Researcher, Unidad Ejecutora Lillo, CONICET, and Professor of Biogeography and Evolution, Universidad Nacional de Tucumán, Tucumán, Argentina.

Professor S. M. Goodman: MacArthur Field Biologist, Field Museum of Natural History, Chicago, Illinois, USA; and Scientific Counselor, Association Vahatra, Antananarivo, Madagascar.

R. Haslauer: Independent Researcher and Member, Zoological Society for the Conservation of Species and Populations (ZGAP), Poernbach, Bavaria, Germany.

S. Hintsche: Biologist for Species Conservation and Mammalogy, Department of Education & Conservation, Hellabrunn Zoo Munich, Munich, Bavaria, Germany.

Professor I. Horáček: Professor, Department of Zoology, Faculty of Sciences, Charles University, Prague, Czech Republic.

A. M. Hutson: Bat Conservation Officer, Fauna and Flora Preservation Society, London; Conservation Officer, Bat Conservation Trust, London; and Freelance Researcher and Consultant, especially on international bat conservation, London, UK.

Professor C. Ibáñez: Professor, Department of Evolutionary Ecology, Estación Biológica de Doñana-CSIC, Sevilla, Spain.

Dr J. Juste: Assistant Professor, Department of Evolutionary Ecology, Estación Biológica de Doñana-CSIC, Sevilla, Spain.

Dr T. Kingston: Professor, Department of Biological Sciences, Texas Tech University, Lubbock, Texas, USA; and Co-Chair, IUCN/SSC Bat Specialist Group.

Dr T. H. Lavery: Honorary Research Fellow, Biodiversity Program, Queensland Museum, Brisbane, Queensland, Australia.

Dr T. E. Lee, Jr: Professor and Chair, Department of Biology, Abilene Christian University, Abilene, Texas, USA.

Dr B. K. Lim: Assistant Curator, Department of Natural History, Royal Ontario Museum, Toronto, Ontario, Canada.

Dr A. López-Baucells: Postdoctoral Researcher, Natural Sciences Museum of Granollers (BiBio - Biodiversity and Bioidicators), Granollers, Catalonia, Spain.

L. O. Loureiro: PhD Candidate, Department of Ecology and Evolutionary Biology, University of Toronto, Toronto, Ontario, Canada.

Professor R. A. Medellín: Professor, Instituto de Ecología, Universidad Nacional Autónoma de México, Ciudad de México, Mexico; and Co-Chair, IUCN/SSC Bat Specialist Group.

Professor A. Monadjem: Professor, Department of Biological Sciences, University of Eswatini, Kwaluseni, Eswatini.

Dr L. M. Moras: Professor, Centro Universitário Metodista Izabela Hendrix, Belo Horizonte, Minas Gerais, Brazil.

Dr R. Moratelli: Research Scientist, Fiocruz Atlantic Forest, Oswaldo Cruz Foundation, Rio de Janeiro, Rio de Janeiro, Brazil.

R. L. M. Novaes: Professor, Department of Biology, Celso Lisboa University, Rio de Janeiro; and PhD Candidate, Federal University of Rio de Janeiro, Rio de Janeiro, Brazil.

B. P. O’Toole: Graduate Student, Department of Biology, Fordham University, Bronx, New York, USA.

Dr A. C. Pavan: Postdoctoral Researcher, Department of Biology, ESALQ/University of São Paulo, Piracicaba, São Paulo, Brazil.

M. Pennay: Project Officer, New South Wales National Parks and Wildlife Service, New South Wales, Australia.

Dr A. Rodríguez-Durán: Professor, Departamento de Ciencias Naturales, Universidad Interamericana, Bayamón, Puerto Rico.

Dr B. Rodríguez-Herrera: Professor and Researcher, Escuela de Biología y Museo de Zoología, Universidad de Costa Rica, San Pedro de Montes de Oca, San José, Costa Rica.

Professor S. J. Rossiter: Head, Department of Biology, School of Biological and Chemical Sciences, Queen Mary University of London, London, UK.

Dr M. Ruedi: Curator, Natural History Museum of Geneva, Geneva, Switzerland.

Dr P. Soisook: Researcher and Curator of Mammals, Princess Maha Chakri Sirindhorn Natural History Museum, Faculty of Science, Prince of Songkla University, Hat Yai, Songkhla, Thailand.

Dr S. Solari: Associate Professor, Instituto de Biología, Universidad de Antioquia, Medellín, Colombia; and Red List Authority, Focal Point New World, IUCN/SSC Bat Specialist Group.

Professor V. da C. Tavares: Professor, Departamento de Sistemática e Ecologia, CCEN/DSE, Universidade Federal da Paraíba, João Pessoa, Paraíba, Brazil.

Professor P. J. Taylor: South African Research Chair on Biodiversity Value and Change, School of Mathematical and Natural Sciences, University of Venda, Thohoyandou, Limpopo, South Africa.

A. Tejedor: Senior Lecturer, The School for Field Studies, Center for Amazon-Andes Studies, Beverly, Massachusetts, USA.

Dr V. D. Thong: Associate Professor, Institute of Ecology and Biological Resources, Graduate University of Science and Technology, Vietnam Academy of Science and Technology, Hanoi, Vietnam.

D. G. Tirira: Scientific Director, Fundación Mamíferos y Conservación, Quito, Ecuador.

Dr C. A. Toth: Postdoctoral Researcher, Department of Biological Sciences, Boise State University, Boise, Idaho, USA.

Dr S. M. Tsang: Research Associate, Department of Mammalogy, American Museum of Natural History, New York, New York, USA; and Research Associate, Zoology Division, National Museum of the Philippines, Manila, Philippines.

Dr M. Tschapka: Associate Professor, Institute of Evolutionary Ecology, University of Ulm, Ulm, Baden-Württemberg, Germany; and Research Associate, Smithsonian Tropical Research Institute, Balboa, Panama.

C. Tuneu-Corral: Research Assistant, Bat Research Group, Natural Sciences Museum of Granollers, Catalonia, Spain.

Dr V. Van Cakenberghe: Research Associate, Functional Morphology, Department of Biology, University of Antwerp, Antwerp, Belgium.

Artists:

Ilian Velikov : (b. Bulgaria, 1985) is a wildlife artist who has had a strong interest in the natural world since an early age. He studied and worked in the animation industry where he mastered the digital drawing and painting techniques which he now uses in his illustrations. Besides drawing wildlife, he is often out in the field observing it and has assisted in a number of mostly herpetological studies. He illustrated the Field Guide to Amphibians & Reptiles of Britain and Europe by J. Speybroeck et al. and since 2018 has worked with Lynx Edicions on a variety of projects including Ocells de Catalunya, País Valencià i Balears, Lynx and BirdLife International Field Guides and the Handbook of the Mammals of the World.

Francesc Jutglar: (b. Barcelona, 1962) is a zoologist with a lifelong vocation for the natural world, and a self-taught scientific illustrator who has worked largely in pencil and watercolour. Long experience of observing birds in the field, and up close while participating in scientific ringing campaigns, has sharpened the precision of his illustrations. His professional career has been closely bound to Lynx Edicions since its foundation. Works of significance include his participation in various Field Guides, notably Ocells de Catalunya, País Valencià i Balears and the first volumes of the Handbook of the Birds of the World. In later volumes and in the series Lynx and Birdlife International Field Guides his role is that of Artwork Co-ordinator. His illustrations have been commissioned by a number of nature parks and for educational publications, and he has imparted courses on nature illustration. Currently his work is accomplished using digital media.

Blanca Martí de Ahumada: (b. Barcelona, 1981) is a scientific and nature illustrator. She graduated from the University of Barcelona in Art History, and is a naturalist by vocation. She specializes in scientific and children’s illustration at the Barcelona school Escola de la Dona de la Diputació and imparts training courses at the Universities of Valencia and Barcelona. Her illustrations have appeared in various national and international publications: scientific and educational books in Catalonia, Costa Rica, Brazil and Nicaragua, and also on posters for natural parks: the Parc Natural del Montnegre i El Corredor, Parc del Foix in metropolitan Barcelona and the Dindefello Reserve in Senegal, among others. She has also illustrated teaching materials for nature resorts and animal reserves such as Molló Parc or Centre d’Estudi i divulgació del Llop Signatus. She is currently working on projects for the publisher Zona Tropical of Costa Rica and participating in the volume Bats from the collection Handbook of the Mammals of the World.

Alex Mascarell Llosa: (b. Barcelona, 1978) is a naturalist, designer and illustrator who works mainly on educational materials and has a special interest in ornithology. In addition to his work designing layouts for a range of teaching materials, he is an animal illustrator. Passionate about birds, he works regularly at Barcelona Zoo, collaborating with many organizations and institutions in the design of materials and the production of original illustrations. He has participated in the Dictionari de las Aus en Català, which collates the Catalan nomenclature of all the bird species of the world, and he works pro bono on drawing projects for other institutions dedicated to the defense of the natural world, both locally and internationally. He has worked on Volume 2 of the Illustrated Checklist of the Birds of the World and on the Birds of Thailand and the Birds of Vietnam from the Lynx and Birdlife International Field Guides Collection.

Faansie Peacock: (b. Pretoria, 1982) is a South African ornithologist, author and artist with a passion for biodiversity and, specifically, sharing the natural world with those around him. Through his various publications, both scientific and popular, Peacock bridges the gap between the academic and public spheres in the hope that increased awareness and affinity to nature ultimately leads to conservation. He specialises in digital painting techniques, which allows him to produce unprecedented numbers of illustrations of exceptional accuracy and quality. Working from his own field sketches together with museum skins and reference photographs, Peacock has authored and illustrated specialist guides on LBJs (larks, pipits, cisticolas, warblers etc.), waders (shorebirds), and threatened species. In line with his current mindset as a father of two young boys, his latest book is a complete field guide to 722 species of South African birds, aimed specifically at children. In addition to books, he has done natural history illustration, design and layout work in a wide variety of formats, from business cards to billboards.

Jesús Rodríguez-Osorio Martín: (b. Madrid, 1965) is a self taught artist. From an early age a strong sense of vocation drew him to the natural world and he used illustration as a way of satisfying his zoological curiosity. Over the years he refined his techniques and in the 1980s published a number of illustrations in Quercus magazine. He illustrated an entire volume of the prestigious Fauna Ibérica (Volume 44: Coleoptera-Hydraenidae). He spent years combining the photography of wild animals with illustration, but with the arrival of digital illustration techniques he opted for these. He is one of the illustrators of Volume 9 of the Handbook of the Mammals of the World.

Lluís Sogorb: (b. Monòver, 1965) is a self-taught painter and illustrator. His abilities and interest in the natural world led him to the illustration and painting of nature. Although he began with traditional techniques like graphite, watercolour, acrylic and oil paint, as soon as digital techniques became available he became interested in them and currently creates almost all his illustrations in this way. His illustrations have appeared in numerous publications and he has taken part in various educational projects centred on the environment and conservation. In recent times he has taken part in various projects involving ornithological tourism, as well as frequently collaborating with different publishers. He has participated in A New European Breeding Bird Atlas, currently in preparation. In 2013 he won the first Concurso de Ilustración de la Naturaleza (Illustration prize) organized by the Sociedad Gaditana de Historia Natural for his image of a Northern Bald Ibis. In 2017 he was awarded a prize in the Painting Animals in the Wild category of the XI Golden Turtle Wildlife Festival (Moscow, Russia) for his painting “The Guardian of the Mountain.”